AZC 46 Supplement (2003)

Issue:

VOL NA

Keywords:

New species, new genus, new family, Diptera, fossil, Triassic, France, phylogeny

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Triassic Diptera: review, revisions and descriptions

• Wiesław KRZEMIŃSKI
• Ewa KRZEMIŃSKA

Abstract:

Triassic Diptera are reviewed and their systematic classification and phylogenetic  importance are discussed. Four new species of Diptera are described from the Lower/Middle Triassic of France: Louisa nova, n. gen. et sp. (Grauvogeliidae), Tanus triassicus, n. gen. et sp. (Nadipteridae), Archilimonia vogesiana, n. gen. et sp. (Archilimoniidae, n. fam.), Vymrhyphus blagoderovi n. sp. (Protorhyphidae) and Gallia alsatica n. gen. et sp. (Rhagionidae), which – together with Grauvogelia arzvilleriana earlier described from the same deposits – represent the oldest Diptera known. The following changes in taxonomy are introduced: family Vladipteridae with species: Vladiptera kovalevi and Dilemmala specula SHCHERBAKOV 1995 is transferred to Mecoptera. The subfamilies: Psychotipinae and Kuperwoodinae are raised to a family rank. Subfamily Gnomuscinae is considered a doubtful Diptera and transferred, with some reservation, to the Trichoceromorpha at the family rank. Family Alinkidae is transferred from infraorder Tabanomorpha to Xylophagomorpha.Two new infraordes are established: Hennigmatomorpha and Tillyardomorpha. Phylogenetic trees of all five suborders of Diptera are presented (i. e., of the Diarchineura, Neoneura, Polyneura, Anisoneura and Brachycera). Representatives of all these suborders are known from the Triassic, and, with exception of the Neoneura, were present already in the Lower/Middle Triassic of France, i. e., the oldest deposits bearing the Diptera.